After the events of the first film, WALL-E (voiced by Ben Burtt) and EVE (voiced by Elissa Knight) have played a crucial role in rebuilding Earth. However, the planet still faces significant challenges. Humanity has returned, but the path to restoring the Earth's ecosystem is far from easy. WALL-E and EVE are sent on a mission to revive nature by planting trees and repairing the devastated soil.

As they traverse the planet, they discover that some lands remain toxic and new creatures, remnants of Earth’s past, have begun to emerge. Their mission becomes a true battle not only against the environment but also against forgotten robots from the past, ready to destroy everything that has been rebuilt.